Polydesmida: Paradoxosomatidae: Australiosomatinae: Australiosomatini: Heterocladosoma species

 

Heterocladosoma asperum (L. Koch, 1867)

  • Heterocladosoma asperum
  • Strongylosoma aspera L. Koch 1867: 245 (first description)
    Koch, L. (1867) Beschreibungen neuer Arachniden und Myriopoden. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien 17: 173-250.
  • Orthomorpha aspera Attems 1898: 328 (new combination)
    Attems, C. (1898) System der Polydesmiden. I. Theil. Denkschriften der kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 67: 221-482, pls 1-11.
  • Strongylosoma asperum Attems 1914: 235 (in species catalogue)
    Attems, C. (1914) Die indo-australischen Myriopoden. Archiv für Naturgeschichte (A) 80(4): 1-398.
  • Orthomorpha aspera Chamberlin 1920: 113 (in species catalogue)
    Chamberlin, R.V. (1920) The Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology 64(1): 1-269.
  • Strongylosoma asperum Jeekel 1968: 19 (in list of Australian species), 27 (in family revision, as species of 'enigmatic' generic status)
    Jeekel, C.A.W. (1968) On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). C.A.W. Jeekel, Rotterdam, 162 pp.
  • 'Strongylosoma asperum' Jeekel 1981: 49 (in list of species recorded from eastern Australia)
    Jeekel, C.A.W. (1981) Australia Expedition 1980; legit C.A.W. Jeekel and A.M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania. Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoölogie (Zoölogisch Museum), Universiteit van Amsterdam 30: 1-59.
  • Heterocladosoma asperum Jeekel 1987: 12 (in list of species recorded from Qld; new combination), 24 (in species key), 28 (redescription); Fig. 9 (p. 30)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma asperum Nguyen and Sierwald 2013: 1170 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Syntype(s): At least one male, Brisbane, Qld, date and collector unknown, NHM [-27.4700 153.0250 ±10 km]
 

Heterocladosoma bifalcatum (Silvestri, 1898)

  • Heterocladosoma bifalcatum
  • Eustrongylosoma bifalcatum Silvestri 1898: 231 (first description); Figs 13, 14 (p. 231)
    Silvestri, F. (1898) Note sui Chilopodi e Diplopodi conservati nel Museo Zoologico di Firenze. I. Alcuni nuovi Diplopodi de Queensland (Cairns). Bullettino della Società Entomologica Italiana 29: 225-232.
  • Australisoma bifalcatum Brölemann 1913: 89 (in list of Australian species, as Eustrongylosoma bifalcatum), 91 (in list of species included in Australiosoma, new combination), 94 (in species key)
    Brölemann, H.W. (1913) The Myriapoda in the Australian Museum. Part ii. Diplopoda. Records of the Australian Museum 10(6): 77-158, pls 14-18.
  • Australiosoma bifalcatum Attems 1914: 225 (in species catalogue)
    Attems, C. (1914) Die indo-australischen Myriopoden. Archiv für Naturgeschichte (A) 80(4): 1-398.
  • Australiosoma bifalcatum Chamberlin 1920: 104 (in species catalogue)
    Chamberlin, R.V. (1920) The Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology 64(1): 1-269.
  • Australiosoma bifalcatum Attems 1937: 234 (in species key), 237 (redescription); Fig. 295 (p. 237)
    Attems, C. (1937) Myriapoda. 3. Polydesmoidea. I. Fam. Strongylosomidae. Pp. i-xxii, 1-300 in Das Tierreich 68. Walter de Gruyter and Co., Berlin.
  • Australiosoma bifalcatus Attems 1944: 249 (distinguished from Australiosoma castaneum, i.e. Oncocladosoma castaneum castaneum)
    Attems, C. (1944) Neue Polydesmoidea. Zoologischer Anzeiger 144: 223-251.
  • Heterocladosoma bifalcatum Jeekel 1968: 17 (in list of Australian species, as Australiosoma bifalcatum), 25 (in family revision, new combination), 144 (as Eustrongylosoma bifalcatum, designated type species of Heterocladosoma); Fig. 2 (p. 145)
    Jeekel, C.A.W. (1968) On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). C.A.W. Jeekel, Rotterdam, 162 pp.
  • Heterocladosoma bifalcatum Jeekel 1981: 47, 49 (in list of species recorded from eastern Australia)
    Jeekel, C.A.W. (1981) Australia Expedition 1980; legit C.A.W. Jeekel and A.M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania. Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoölogie (Zoölogisch Museum), Universiteit van Amsterdam 30: 1-59.
  • Heterocladosoma bifalcatum Jeekel 1985: 19 (as Eustrongylosoma bifalcatum, cited as type species of Heterocladosoma), 22 (in taxonomic discussion)
    Jeekel, C.A.W. (1985) Millipedes from Australia, 5: Australiosomatini from South Australia, with a note on the status of Polydesmus innotatus Karsch, and first record of a second Mediterranean julid in Australia (Diplopoda: Polydesmida, Paradoxosomatidae and Julida, Julidae). Records of the South Australian Museum 19(3): 19-37.
  • Heterocladosoma bifalcatum Jeekel 1987: 12 (in list of species recorded from Qld), 24 (in species key, and redescription); Figs 5-8 (p. 26)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Eustrongylosoma bifalcatum Shelley et al. 2000: 105 (in genus catalogue, as type of Heterocladosoma)
    Shelley, R.M., Sierwald, P., Kiser, S.B. and Golovatch, S.I. (2000) Nomenclator generum et familiarum Diplopodorum II. A List of the Genus and Family-Group Names in the Class Diplopoda from 1958 through 1999. Pensoft, Sofia, 167 pp.
  • Heterocladosoma bifalcatum Rowe and Sierwald 2006: 541 (as Eustrongylosoma bifalcatum, type species of Heterocladosoma; as Australiosoma bifalcatum, in synonymy; redescription); Figs 4 (p. 532), 10 (p. 535), 26, 27 (p. 545), 38, 39 (p. 548), 47, 48 (p. 550)
    Rowe, M. and Sierwald, P. (2006) Morphological and systematic study of the tribe Australiosomatini (Diplopoda: Polydesmida: Paradoxosomatidea: Paradoxosomatidae) and a revision of the genus Australiosoma Brölemann. Invertebrate Systematics 20(5): 527-556.
  • Heterocladosoma bifalcatum Nguyen and Sierwald 2013: 1170 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Syntype(s): At least one male, Cairns, Qld, December 1891, [Giovanni] Podenzana, MSNF [-16.9250 145.7740 ±5 km]
  • Note: Jeekel (1987: 28) believes the Cairns locality is in error and recorded it from Colosseum, Qld (Jeekel 1987: 24). This species is not only abundant in the Brisbane area but is established as far south as Sydney (see the Localities section of this website).
 

Heterocladosoma galaxias Jeekel, 1985

  • Heterocladosoma galaxias
  • Heterocladosoma galaxias Jeekel 1985: 22 (first description); Fig. 2 (p. 23)
    Jeekel, C.A.W. (1985) Millipedes from Australia, 5: Australiosomatini from South Australia, with a note on the status of Polydesmus innotatus Karsch, and first record of a second Mediterranean julid in Australia (Diplopoda: Polydesmida, Paradoxosomatidae and Julida, Julidae). Records of the South Australian Museum 19(3): 19-37.
  • Heterocladosoma galaxias Jeekel 1987: 24 (in species key)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma galaxias Jeekel 2002: 77 (in list of species recorded from SA)
    Jeekel, C.A.W. (2002) Millipedes from Australia, 14: A third contribution to the South Australian Paradoxosomatidae (Diplopoda, Polydesmida). Myriapod Memoranda 5: 60-77.
  • Heterocladosoma galaxias Nguyen and Sierwald 2013: 1170 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Holotype: Male, Gammon Plateau, SA, 19 September 1956, G.F. Gross, SAM OM1336 [-30.5333 138.5500 ±50 km!!]
  • Paratype(s): None
 

Heterocladosoma hamuligerum (Verhoeff, 1924)

  • Heterocladosoma hamuligerum
  • Australiosoma hamuligerum Verhoeff 1924: 13 (in taxonomic discussion), 24 (in species key), 26 (first description); Fig. 15, pl. 1
    Verhoeff, K.W. (1924) Results of Dr. E. Mjöberg's scientific expeditions to Australia 1910-1913. 34. Myriapoda: Diplopoda. Arkiv för Zoologi 16(5): 1-142, pls 1-5.
  • Australiosoma hamuligerum Verhoeff 1928: 88 (in taxonomic discussion)
    Verhoeff, K.W. (1928) On Diplopoda in the Australian Museum, Sydney. Records of the Australian Museum 16(2): 79-115, pls 6-12.
  • Australiosoma hamuligerum Attems 1937: 234 (in species key), 236 (redescription)
    Attems, C. (1937) Myriapoda. 3. Polydesmoidea. I. Fam. Strongylosomidae. Pp. i-xxii, 1-300 in Das Tierreich 68. Walter de Gruyter and Co., Berlin.
  • Heterocladosoma hamuligerum Jeekel 1968: 17 (in list of Australian species, as Australiosoma hamuligerum), 25 (in family revision, new combination), 144 (as Australiosoma hamuligerum, included in new genus Heterocladosoma)
    Jeekel, C.A.W. (1968) On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). C.A.W. Jeekel, Rotterdam, 162 pp.
  • Heterocladosoma hamuligerum Jeekel 1981: 49 (in list of species recorded from eastern Australia)
    Jeekel, C.A.W. (1981) Australia Expedition 1980; legit C.A.W. Jeekel and A.M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania. Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoölogie (Zoölogisch Museum), Universiteit van Amsterdam 30: 1-59.
  • Heterocladosoma hamuligerum Jeekel 1985: 22 (in taxonomic discussion)
    Jeekel, C.A.W. (1985) Millipedes from Australia, 5: Australiosomatini from South Australia, with a note on the status of Polydesmus innotatus Karsch, and first record of a second Mediterranean julid in Australia (Diplopoda: Polydesmida, Paradoxosomatidae and Julida, Julidae). Records of the South Australian Museum 19(3): 19-37.
  • Heterocladosoma hamuligerum Jeekel 1987: 12 (in list of species recorded from Qld), 24 (in species key), 32 (redescription); Figs 14, 15 (p. 36)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma hamuligerum Nguyen and Sierwald 2013: 1171 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Holotype: Male, Blackall Range, Qld, 19 September 1911, E. Mjöberg, possibly in NHRM [-26.6870 152.8520 ±5 km]
  • Paratype(s): None
 

Heterocladosoma trabeatum Jeekel, 1987

  • Heterocladosoma trabeatum
  • Heterocladosoma trabeatum Jeekel 1987: 12 (in list of species recorded from Qld), 24 (in species key), 28 (first description); Fig. 10 (p. 30)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma trabeatum Nguyen and Sierwald 2013: 1171 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Holotype: Male, [?Brisbane] 'Vicinity of Priceden', Qld, Eucalyptus forest, under Eucalyptus bark, 28 February 1977, G.F. Kurceva, ZMMU
  • Paratype(s): None
 

Heterocladosoma transversetaeniatum perarmatum Jeekel, 1987

  • Heterocladosoma transversetaeniatum perarmatum
  • Heterocladosoma transversetaeniatum perarmatum Jeekel 1987: 12 (in list of species recorded from Qld), 31 (first description); Figs. 12, 13 (p.33)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma transversetaeniatum perarmatum Nguyen and Sierwald 2013: 1171 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Holotype: Male, Rockhampton, Qld, 15 November 1924, C. Barrett, AMNH [-23.3770 150.5100 ±5 km]
  • Paratype(s): 2 males, 1 female, details as for holotype, AMNH
 

Heterocladosoma transversetaeniatum transversetaeniatum (L. Koch, 1867)

  • Heterocladosoma transversetaeniatum transversetaeniatum
  • Strongylosoma transverse-taeniatum L. Koch 1867: 246 (first description)
    Koch, L. (1867) Beschreibungen neuer Arachniden und Myriopoden. Verhandlungen der kaiserlich-königlichen zoologisch-botanischen Gesellschaft in Wien 17: 173-250.
  • Polydesmus (Strongylosoma) transversetaeniatum Karsch 1881: 44 (in species list); Fig. 23, pl. 3
    Karsch, F. (1881) Zum Studium der Myriopoda Polydesmia. Archiv für Naturgeschichte 47(1): 36-49, pl. 3.
  • Strongylosoma transversetaeniatum Attems 1898: 281 (in species key), 306 (redescription); Figs 18, 19, pl. 1
    Attems, C. (1898) System der Polydesmiden. I. Theil. Denkschriften der kaiserlichen Akademie der Wissenschaften. Mathematisch-Naturwissenschaftliche Klasse 67: 221-482, pls 1-11.
  • Australiosoma transverse-taeniatum Brölemann 1913: 89 (in list of Australian species, as Polydesmus (Strongylosoma) transverse-taeniatum), 91 (in list of species included in Australiosoma, new combination), 94 (in species key)
    Brölemann, H.W. (1913) The Myriapoda in the Australian Museum. Part ii. Diplopoda. Records of the Australian Museum 10(6): 77-158, pls 14-18.
  • Australiosoma transversetaeniatum Brölemann 1916: 539 (in taxonomic discussion)
    Brölemann, H.W. (1916) Essai de classification des Polydesmiens (Myriapodes). Annales de la Société Entomologique de France 84: 523-608.
  • Australiosoma transversetaeniatum Attems 1914: 225 (in species catalogue)
    Attems, C. (1914) Die indo-australischen Myriopoden. Archiv für Naturgeschichte (A) 80(4): 1-398.
  • Australiosoma transversetaeniatum Chamberlin 1920: 104 (in species catalogue)
    Chamberlin, R.V. (1920) The Myriopoda of the Australian region. Bulletin of the Museum of Comparative Zoology 64(1): 1-269.
  • Australiosoma transversetaeniatum Attems 1937: 234 (in species key), 237 (redescription); Fig. 296 (p. 238)
    Attems, C. (1937) Myriapoda. 3. Polydesmoidea. I. Fam. Strongylosomidae. Pp. i-xxii, 1-300 in Das Tierreich 68. Walter de Gruyter and Co., Berlin.
  • Heterocladosoma transversetaeniatum Jeekel 1968: 17 (in list of Australian species, as Australiosoma transversetaeniatum), 25 (in family revision, new combination), 144 (as Strongylosoma transversetaeniatum, included in new genus Heterocladosoma)
    Jeekel, C.A.W. (1968) On the Classification and Geographical Distribution of the Family Paradoxosomatidae (Diplopoda, Polydesmida). C.A.W. Jeekel, Rotterdam, 162 pp.
  • Heterocladosoma transversetaeniatum Jeekel 1981: 49 (in list of species recorded from eastern Australia)
    Jeekel, C.A.W. (1981) Australia Expedition 1980; legit C.A.W. Jeekel and A.M. Jeekel-Rijvers. List of collecting stations, together with general notes on the distribution of millipedes in eastern Australia and Tasmania. Verslagen en Technische Gegevens, Instituut voor Taxonomische Zoölogie (Zoölogisch Museum), Universiteit van Amsterdam 30: 1-59.
  • Heterocladosoma transversetaeniatum Jeekel 1985: 22 (in taxonomic discussion)
    Jeekel, C.A.W. (1985) Millipedes from Australia, 5: Australiosomatini from South Australia, with a note on the status of Polydesmus innotatus Karsch, and first record of a second Mediterranean julid in Australia (Diplopoda: Polydesmida, Paradoxosomatidae and Julida, Julidae). Records of the South Australian Museum 19(3): 19-37.
  • Heterocladosoma transversetaeniatum Jeekel 1987: 12 (in list of species recorded from Qld), 31 (in taxonomic discussion)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma transversetaeniatum transversetaeniatum Nguyen and Sierwald 2013: 1171 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Note: The types situation is unclear. Karsch (1881b: 44) says ZMB holds specimens from Cape York (collected by 'Salmin') and Sydney (collected by 'Daemel'). Jeekel (1987) illustrates the right gonopod of 'the male holotype in the Hamburg Museum' (p. 31), from Brisbane, Qld (-27.4700 153.0250 ± 10 km), as drawn by R.L. Hoffman.
 

Heterocladosoma zebratum Jeekel, 1985

  • Heterocladosoma zebratum
  • Heterocladosoma zebratum Jeekel 1985: 20 (first description); Fig. 1 (p. 21)
    Jeekel, C.A.W. (1985) Millipedes from Australia, 5: Australiosomatini from South Australia, with a note on the status of Polydesmus innotatus Karsch, and first record of a second Mediterranean julid in Australia (Diplopoda: Polydesmida, Paradoxosomatidae and Julida, Julidae). Records of the South Australian Museum 19(3): 19-37.
  • Heterocladosoma zebratum Jeekel 1987: 24 (in species key)
    Jeekel, C.A.W. (1987) Millipedes from Australia, 11: Australiosomatini from Queensland (Diplopoda, Polydesmida, Paradoxosomatidae). Beaufortia 37(2): 11-41.
  • Heterocladosoma zebratum Jeekel 2002: 65 (partial redescription), 77 (in list of species recorded from SA); Fig. 4 (p. 62)
    Jeekel, C.A.W. (2002) Millipedes from Australia, 14: A third contribution to the South Australian Paradoxosomatidae (Diplopoda, Polydesmida). Myriapod Memoranda 5: 60-77.
  • Heterocladosoma zebratum Nguyen and Sierwald 2013: 1171 (in Paradoxosomatidae catalogue)
    Nguyen, A.D. and Sierwald, P. (2013) A worldwide catalog of the family Paradoxosomatidae Daday, 1889 (Diplopoda: Polydesmida). Check List 9(6): 1132-1353.
  • Holotype: Male, Coralbignie to Buckleboo, SA, 15-16 March 1950, G.F. Gross, SAM sample missing [-32.7667 136.2667 ±5 km (halfway)]
  • Paratype(s): 3 females, details as for holotype, SAM OM1327, OM1328
  • Paratype(s): 1 male, N of Kokotha, SA, 11 June 1956, G.F. Gross, SAM sample missing [-31.2500 135.2333 ±10 km]
  • Paratype(s): 1 male, Nonning, Gawler Ranges, SA, burnt out of spinifex, 17 June 1956, G.F. Gross, SAM OM1392 [-32.5167 136.4833 ±10 km]